RESEARCH GRANTS
National Institutes of Health Academic
Research Enhancement Award (AREA Grant, 1999-2002).
Title: "Galanin Regulation of Pituitary
Reproductive Hormones"
Amount of award: $74,222.
Summary of proposed research activities:
|
A
substantial body of evidence now exists which implicates a role for the
hypothalamic neuropeptide galanin in the regulation of
pituitary reproductive hormone secretion. Galanin neurons regulate
pituitary luteinizing hormone (LH) secretion at two distinct levels: 1).
at the level of the hypothalamus, where galanin indirectly
regulates LH by stimulating the production of the hypothalamic releasing
factor gonadotropin-releasing hormone (GnRH), and 2). at the level of the
pituitary gland, where galanin directly stimulates the
secretion of LH from gonadotropes and enhances GnRH-stimulated LH
secretion. While the hypothalamic effects of galanin on GnRH secretion are
relatively well-characterized, little is known about galanin regulation of
LH secretion directly at the level of the pituitary gland. Experiments
outlined in this proposal were designed to characterize the pituitary
effects of galanin on reproductive hormone secretion in both males and
females, using the rat as an animal model. Specific aim #1 of this
proposal is to determine whether gonadal steroids alter gonadotrope
sensitivity to galanin stimulation. This hypothesis will be tested in
a series of in vivo infusion experiments examining LH secretory
responses to galanin in 1) intact, castrated, and castrated +
testosterone-replaced male rats; and 2) female rats at different
timepoints during the four day estrous cycle. Studies in females will
specifically address whether gonadotrope sensitivity to galanin
stimulation is upregulated within the context of the proestrous (i.e.
preovulatory) endocrine milieu. Experiments of Specific aim #2 are
designed to characterize the galanin receptor subtype that is expressed
in gonadotropes in the anterior pituitary gland. Double-label in
situ hybridization experiments will be performed to determine if
galanin type 2 receptor (GalR2) messenger RNA (mRNA) expression is present
in anterior pituitary cells that respond to GnRH, and if so, to determine
the relative levels of GalR2 mRNA expression in gonadotropes vs. other
anterior pituitary cell types that respond to galanin. Specific Aim #3 of
this proposal is to investigate the effects of galanin on reproductive
hormone gene expression in the anterior pituitary gland. An in
vitro perifusion system will be used to determine if galanin, in
addition to stimulating the secretion of LH, also has an effect on
the mRNA expression of the LHb
subunit. Furthermore, the effects of galanin on GnRH receptor mRNA
expression will be assessed. Results from these studies will provide
important information regarding the role of galanin in regulating LH
synthesis and secretion from pituitary gonadotropes, events that are
critically important for maintaining reproductive competence in both males
and females. Additionally, participation in the execution of experiments
outlined in this proposal will provide a unique opportunity for
undergraduate students to learn fundamental techniques in reproductive
physiology and molecular biology, experiences that will
serve them well as they pursue their graduate studies. |
National Science Foundation Course, Curriculum
and Laboratory Improvement Grant
(CCLI Grant; 1999-2001).
Title of Grant : "Improvement of Molecular
Biology Components of the Human Biology Curriculum: Incorporation of a
Research-Based Learning Approach".
Amount of award: $44,701 from NSF, with a match
of $44,701 from UW-Green Bay.
Summary of proposed research activities:
The Biology and Human Biology Programs at the
University of Wisconsin (UW)- Green Bay wish to incorporate
modern methods of molecular and cell biology into the undergraduate
curriculum. Specifically, funds were obtained to purchase equipment that will
allow students to perform in situ analyses of protein and/or gene
expression. The Developmental Biology Laboratory Course will utilize the
equipment to conduct laboratory exercises that utilize a research project
approach in order to assess whether this learning approach enhances
student scientific reasoning and analytical skills. Developmental Biology
students will design experiments to analyze the tissue-specific expression of a
gene during embryogenesis, perform labeling experiments for visualization of
this expression in embryo sections generated with the requested cryostats,
record and analyze their data using the requested image analysis station, and
disseminate their scientific data at a student poster session at the end of the
semester and possibly at a UW - System Student Research Conference at the end of
the academic year. Cell Biology students will perform a similar laboratory
project involving enzyme cytochemistry. Finally, other courses (Molecular
Biology, Immunology, Microscopic Anatomy and Molecular Genetics) as well as a
students conducting independent research projects will utilize the requested
equipment for performing related laboratory projects or learning activities
during the course of the academic year. The implementation of these techniques
into the Biology and Human Biology curricula will provide students with
sophisticated training in modern methods of molecular / cell biology, including
tissue cryosectioning, immunocytochemistry, enzyme cytochemistry and in situ
hybridization for the analysis of gene expression. Furthermore, while conducting
these methods within the context of a research project and not a
"canned" laboratory exercise, it is anticipated that student
scientific reasoning / analytical skills will be sharpened.